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Upper Mississippi and Missouri River drainage areas from the eastern slopes of the Rocky Mountains east to Lake Michigan, and from southern Manitoba and probably southwestern Ontario and southeastern Alberta south to southern Oklahoma and Arkansas.

The smallest specimens live in southern Florida, while those near the northern limits of the raccoon's range tend to be the largest (see Bergmann's rule).

The original habitats of the raccoon are deciduous and mixed forests, but due to their adaptability they have extended their range to mountainous areas, coastal marshes, and urban areas, where some homeowners consider them to be pests.

As a result of escapes and deliberate introductions in the mid-20th century, raccoons are now also distributed across mainland Europe, Caucasia, and Japan.

A fifth island raccoon population, the Cozumel raccoon, which weighs only 3 to 4 kg (6.6 to 8.8 lb) and has notably small teeth, is still regarded as a separate species.

The four smallest raccoon subspecies, with a typical weight of 1.8 to 2.7 kg (4.0 to 6.0 lb), live along the southern coast of Florida and on the adjacent islands; an example is the Ten Thousand Islands raccoon (Procyon lotor marinus).

Unlike other procyonids, such as the crab-eating raccoon (Procyon cancrivorus), the ancestors of the common raccoon left tropical and subtropical areas and migrated farther north about 2.5 million years ago, in a migration that has been confirmed by the discovery of fossils in the Great Plains dating back to the middle of the Pliocene.

Four of these subspecies living only on small Central American and Caribbean islands were often regarded as distinct species after their discovery.

It was recorded on John Smith's list of Powhatan words as aroughcun, and on that of William Strachey as arathkone.

In the 1830s, the United States Whig Party used the raccoon as an emblem, causing them to be pejoratively known as 'coons' by their political opponents, who saw them as too sympathetic to African-Americans.

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The two most widespread subspecies are the eastern raccoon (Procyon lotor lotor) and the Upper Mississippi Valley raccoon (Procyon lotor hirtus). states and Canadian provinces to the north of South Carolina and Tennessee.

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